Agoudim Formation

Agoudim Formation
Stratigraphic range: Late Toarcian–Bajocian
Agoudim village
TypeGeological Formation
Underlies
  • R'cifa Formation
Overlies
Thickness2,000 m (6,600 ft) (up to 5,000 m (16,000 ft) reported)
Lithology
PrimaryMarl, Marly Limestone, Bioclastic limestone
Location
LocationEr-Rich
Coordinates32°15′N 4°30′W / 32.25°N 4.50°W / 32.25; -4.50
Approximate paleocoordinates27°00′N 3°00′W / 27.0°N 3.0°W / 27.0; -3.0
RegionHigh Atlas and Middle Atlas
CountryMorocco
Type section
Named forAgoudim
Named byM. Studer
Year defined1980
RegionEr-Rich, Midelt Province
Thickness at type section~4,500–5,000 m (14,800–16,400 ft)
Agoudim Formation (Morocco)

The Agoudim Formation (also known as the "Marnes et Marno-Calcaires d'Agoudim") is a geological formation of late Toarcian to Bajocian (Lower–Middle Jurassic[1]) age in the central High Atlas and Middle Atlas (particularly around Rich and extending to areas like Midelt, Boulemane, and Beni Mellal) of Morocco.[2][3] The formation is also known as the "Formation Marneuse d’Amane-Ilila", "Marnes de Boulemane", "Marnes de Taffert" and "Calcaires à Cancellophycus".[2][4]

Description

The Agoudim Formation is a substantial geological unit primarily composed of marls and marly limestones, representing a continuation from the underlying Tagoudite or Jbel Ouchbis formations and marking a significant shift toward more clay-rich (argillaceous) sedimentation.[2][5] This shift is periodically interrupted by layers of bioclastic material (deposits rich in shell fragments and other organic debris) that increase in both frequency and thickness moving upward through the formation. It is stratigraphically divided into two main members: a lower marly member dominated by gray-blue marls with occasional nodular bioclastic limestone beds, and an upper member characterized by more calcareous and bioclastic rocks with greater lithological variability.[6] The boundary between these members is often marked by a distinctive layer of organogenic turbidites, notably the "Calcaires à Cancellophycus", which forms a prominent brown cuesta and serves as an important marker horizon for geological mapping.[5][7]

The lower member consists of monotonous, micritic marls that reflect relatively quiet hemipelagic depositional conditions, where fine clay and carbonate mud settled along with benthic material.[2] This distal sedimentation was influenced by intermittent turbiditic currents transporting detrital quartz and glauconite, indicating periodic sediment input from nearby land areas and submarine slopes. In contrast, the upper member shows increasingly energetic conditions with cycles of sedimentation starting with lumachellic limestones (shell-rich limestones with oblique lamination) and ending in thick bioclastic marls.[5][6] This member records frequent synsedimentary slumps and spectacular deformation structures like ball-and-pillow folds (phacoids), which are indicative of tectonic instability and sediment loading on unstable slopes. The evolution upward culminates with a thick limestone cap dominated by oolitic limestones and bioherms (reef-like carbonate buildups that further highlight the transition to shallower marine conditions).[6][7]

Paleoenvironment

The Agoudim Formation records multiple Paleoenvironments in a Shallowing basin, from Cold Seep deposits to shallower Reefs

The Agoudim Formation records a progressive transition from deep to shallow marine settings under the influence of rifting, subsidence, and environmental perturbations in a Diapiric Rift Basin.[8][9] The lower marly members were deposited in hemipelagic conditions below storm wave base, with pelagic sedimentation, fine siliciclastic influx from nearby uplifts, and reworked carbonate from Liassic platforms.[10][11] Fossils such as ammonites, radiolarians, foraminifera, small bivalves, and echinoderm debris confirm a low-energy offshore environment. Rapid subsidence and sea-level rise during the Early Bajocian led to a drowning phase of the neritic carbonate factory, favoring marl sedimentation locally enriched in organic matter.[12] Within these successions, several intervals of authigenic carbonates developed, notably yellow micritic nodules, tubular and chimney-like concretions, and conical chemoherms, interpreted as the record of hydrocarbon Cold seep. Petrographic, isotopic, and sedimentological evidence indicates that the second and third authigenic horizons formed at active seep sites, with feeder-pipes, carbonate crusts, microbialites, and bioclast-rich infills, while the yellow nodules of the lower interval likely reflect organic matter remineralization without direct seepage.[13] Strongly depleted Δ13C values (down to −19‰) point to fluids rich in dissolved inorganic carbon generated by organic matter degradation, likely through bacterial sulfate reduction and anaerobic oxidation of hydrocarbons, rather than pure methane seepage. These processes created localized hardgrounds, serpulidae and oyster colonization, and chemoherm buildups on the seafloor.[13][14] As sedimentation continued, carbonate production recovered, and three reefal complexes developed, ranging from microbial-coral patch reefs to larger coral–sponge–algal bioherms, accompanied by ooid–peloid shoals and brachiopod-rich limestones.[15][16][17] These buildups evolved from small mounds to coalescing reefs, shaped by constructive growth, storm reworking, bioerosion, and micritization.[15][16][17] The upper part of the formation is dominated by shallow platform deposits with abundant brachiopods, gastropods, corals, and reefal frameworks.[15][17][18][19] Some areas, like the Agoudim valley record excepcionally preserved platform sections, showing it´s inner ramp had high-energy ooid–peloid shoals, microbial mounds, and small patch reefs and a proximal middle ramp with mixed coral–sponge–bryozoan–brachiopod assemblages in lower energy waters.[20] The distal ramp consisted of marl–limestone alternations with storm beds and oyster–coral–sponge buildups. Local syndepositional highs and terrigenous input created semi-restricted zones with fluctuating oxygenation and nutrients.[20]

Biota

Foranimifera

Taxa Species Locality Member Material Notes Images
Ammobaculites[21][22] fontinensis Talghemt Ridge II-IV Tests Lituolidae
agglutinans Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Lituolidae
Bullopora[13] sp. Jebel Bou Kendill III Tests Polymorphinidae
Citharina[21] serratocostata Talghemt Ridge IV Tests Vaginulinidae
Citharina[21] sp. Talghemt Ridge I-II Tests Vaginulinidae
Eoguttulina[21][22] bilocularis Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Polymorphinidae
sp. Talghemt Ridge IV Tests Polymorphinidae
Epistommina[22] mosquensis Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II-IV Tests Epistomininae
Flabellammina[21][22] althoffi Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge III-IV Tests Lituolidae
Garantella[22] ampasindavaensis Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Epistomininae
stellata Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Epistomininae
sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi II Tests Epistomininae
Haplophragmium[21][22] subaequale Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge III-IV Tests Lituolidae
aequale Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Lituolidae
Ichthyolaria[21] carinata Talghemt Ridge I Tests Ichthyolariidae
gr. major Talghemt Ridge I Tests Ichthyolariidae
sp. Talghemt Ridge I Tests Ichthyolariidae
Lenticulina[21][22] argonauta Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge IV Tests Nodosariidae
chicheryi Talghemt Ridge I-II Tests Nodosariidae
d'orbignyi Talghemt Ridge I-II Tests Nodosariidae
münsteri Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge I-II Tests Nodosariidae
quenstedti Talghemt Ridge IV Tests Nodosariidae
subalata Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge I-II Tests Nodosariidae
toarcense Talghemt Ridge I Tests Nodosariidae
polymorpha Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Nodosariidae
Parurgonina[22] sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Parurgoninidae
Planularia[21] sp. Talghemt Ridge IV Tests Vaginulinidae
Proteonina[21] gr. difflugiformis Talghemt Ridge IV Tests Hormosinidae
Protoglobigerina?[21] spp. Talghemt Ridge I-III Tests Globigerinidae
Pseudonodosaria[21] gr. pygmea Talghemt Ridge I Tests Nodosariidae
multicostata Talghemt Ridge I Tests Nodosariidae
Reinholdella[22] dreheri Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi III-IV Tests Ceratobuliminidae
Spirillina[21] orbicula Talghemt Ridge I-II Tests Spirillinidae
Textularia[21] jurassica Talghemt Ridge IV Tests Textulariidae
Trochammina[21] globigeriniformis Talghemt Ridge IV Tests Trochamminidae
Verneuilinoides[21] cf. subvitreus Talghemt Ridge I-II Tests Verneuilinidae

Ichnotaxa

Ichnogenus Ichnospecies Locality Member Made by Images
Chondrites[23] isp. Assameur, Jebel Taferdout III-IV
Planolites[23] isp. Assameur, Jebel Taferdout III-IV
  • Annelids
  • Bivalves
  • Arthropods
 .
Phycodes[23] isp. Assameur, Jebel Taferdout III-IV
Thalassinoides[13][24] isp. Jebel Bou Kendill III
Trypanites[23] isp. Assameur, Jebel Taferdout III-IV
Zoophycos[25] (Cancellophycus) ispp. Amellago; Talghemt Ridge; Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi I-IV

Brachiopoda

Taxa Species Locality Member Material Notes Images
Acanthothiris[26] oligocantha Rich (syncline of Jebel Bou Kendill) II Shells Athyrididae
tenuispina Rich (syncline of Jebel Bou Kendill) III Shells Athyrididae
Cymatorhynchia[27] quadriplicata Jebel Assameur; Boukendil; Rich-Gourrama area IV Shells Tetrarhynchiidae
Kallirhynchia[28] concinna Amellago, Talghemt Ridge I Shells Rhynchonellidae
Loboidothyris[26] sp. Rich (syncline of Jebel Bou Kendill) II Shells Loboidothyrididae
Lophrothyris[26] contracta Rich (syncline of Jebel Bou Kendill) II Shells Terebratulidae
Monsardithyris[17][26] cortonensis Rich (syncline of Jebel Bou Kendill) III Shells Terebratulidae
ventricosa Assameur n’Ait Fergane IV Shells Terebratulidae
sp. cf. M. loubensis Bou-Ouchène (northern platform); Rich III Shells Terebratulidae
Quadratirhynchia[28] vasconcellosi Amellago I Shells Tetrarhynchiidae
Rhactorhynchia[26] distracta Rich (syncline of Jebel Bou Kendill) III Shells Rhynchonellidae
Rhynchonelloidea[26] goyi Amellago I Shells Rhynchonellidae
Rugitela[26] cf. hughesi Rich (syncline of Jebel Bou Kendill) II–III Shells Zeilleriidae
Zeilleria[17][26] truncatella Rich (syncline of Jebel Bou Kendill) III Shells Zeilleriidae
perlata Assameur n’Ait Fergane, Boukendil IV Shells Zeilleriidae

Bryozoa

Genus Species Locality Member Material Notes Images
Berenicea[17] sp. Boukendil, Assameur IV Colonies Bereniceidae
Collapora[17] cf. straminea Boukendil IV Colonies Atactotoechidae
Entalophora[17] cf. stokesi Boukendil IV Colonies Entalophoridae

Cnidaria

Genus Species Locality Member Material Notes Images
Acropora[17] cf.sp. Assameur n’Ait Fergane, Boukendil IV Colonies Acroporidae
Aggomorphastraea[29] "sp. 1" Amellago I Colonies Latomeandridae
Allocoenia[29] "sp. 1" Amellago I Colonies Actinastreidae
"sp. 2" Amellago I Colonies Actinastreidae
Ampakabastrea[29] ampakabensis Amellago I Colonies Astrocoeniidae
Cladophyllia[9][17] sp. Assameur n’Ait Fergane; Imilchil I-IV Colonies Caryophylliidae
Comophyllia[17] sp. Assameur n’Ait Fergane IV Colonies Comophylliidae
Complexastrea[29] "sp. 1" Amellago I Colonies Thecosmiliidae
Cyathocoenia[29] faugeresi Amellago I Colonies Actinastreidae
"sp. 1" Amellago I Colonies Actinastreidae
Dendrarea[17] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Dasmosmilia[29] "sp. 1" Amellago I Colonies Dermosmiliidae
Dimorpharaea[17] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Edwardoseris[17] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Enallocoenia[29] "sp. 1" Amellago I Colonies Actinastreidae
Isastrea[17][29] sp. Assameur n’Ait Fergane IV Colonies Latomeandridae/Montlivaltiidae
"sp. 1" Amellago I Colonies Latomeandridae
Kobymeandra[17] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Latiastrea[29] "sp. 1" Amellago I Colonies Latomeandridae
Lochmaeosmilia[9] sp. Imilchil I Colonies Calamophylliidae
Microphyllia[29] irregularis Amellago I Colonies Latomeandridae
Montlivaltia[17] sp. Assameur n’Ait Fergane IV Coralla Montlivaltiidae (solitary)
Polystylidium[17] sp. Assameur n’Ait Fergane IV Colonies Scleractinia (family uncertain)
Stylosmilia[17][29] sp. Foum Zabel (southern margin) II-III Colonies Thecosmiliidae
"sp. 1" Amellago I Colonies Thecosmiliidae
Thecocyathus[29] mactrus Amellago I Solitary corallites Caryophylliidae
Vallimeandropsis[29] lineata Amellago I Colonies Scleractinia (family uncertain)

Porifera

Genus Species Locality Member Material Notes Images
Cheilosporites[23] tirolensis Assameur, Jebel Taferdout IV Sponge body fossils Vaceletidan
spp. Assameur, Jebel Taferdout IV Sponge body fossils
Cladocoropsis[17] spp. Boukendil, Assameur IV Sponge body fossils Axinellidan
Eudea[17] cf. globata Boukendil, Assameur IV Sponge body fossils Stellispongiida
Neuropora[9] spp. Imilchil I Sponge body fossils Neuroporidae

Mollusks

Taxa Species Locality Member Material Notes Images
Actinostreon[13][16] sp. Assameur n’Ait Fergane, Boukendil IV Shells Gryphaeidae
Alocolytoceras[1][30] ophioneum Gheris Valley I Shells Lytoceratidae
dorcadis Ait El Abbas; Azag I Shells
sp. Azag I Shells
Ambersites[1] sp. Rich region II–III Shells Hammatoceratidae
Amphitrochus[31] abbas Rich region; Outarbat III-IV Shells Nododelphinulidae
Ancolioceras[32] opalinoides n’Zala (Jbel Aouja) I–II Shells Graphoceratidae
Asthenoceras[1] sp. Rich (Concavum–Discites zones) II–III Shells Sonniniidae
Belemnopsis[33] baculiformis Iwansitn III Rostra Belemnopseidae
pougatchevskae Iwansitn III Rostra
Bositra[1] buchi Amellago; Rich I–III Shells Posidoniidae
Botula[16] sp. Boukendil area IV Shells, borings Mytilidae
Bradfordia[1] costata Rich region III Shells Oppeliidae
sp. Rich region III Shells
Calabribelus[33] combemoreli Rich (Jbel Bou Kandil), Iwansitn (Middle Atlas) II–III Rostra Holcobelidae
tetramerus Rich, Iwansitn II–III Rostra
Calliphylloceras[30][34] nilssoni Amellago I Shells Phylloceratidae
cf.capitanii Azag I-II Shells
sp. Azag I-II Shells
Catacoeloceras[34] sp. Ait El Abbas I Shells Dactylioceratidae
Catulloceras[1] sp. n’Zala (Jbel Aouja) I Shells Graphoceratidae
Chlamys[1][9] sp. Rich, Amellago; Imilchil I-II–III Shells Pectinidae
Cirrus[31] sp. Aberdouz III-IV Shells Cirridae
Conobelemnopsis[33] sp. A Rich, Iwansitn III Rostra Belemnopseidae
Collina[30] gemma Ait El Abbas; Azag I Shells Dactylioceratidae
cf.nummularia Ait El Abbas I Shells
linae Ait El Abbas I Shells
sp. Azag I Shells
Cotteswoldia[32] sp. n’Zala (Jbel Aouja) I Shells Graphoceratidae
Crassiceras[30] cf.latum Ait El Abbas I Shells Hildoceratidae
Ctenostreon[1] sp. Rich, Amellago II–III Shells Limidae
Cypholioceras[32] sp. n’Zala (Jbel Aouja) II Shells Graphoceratidae
Dactylioceras[21][30][34] (Eodactylites) mirabile Amellago I Shells Dactylioceratidae
(Eodactylites) pseudocommune Amellago I Shells
(Eodactylites) simplex Talghemt Ridge I Shells
(Orthodactylites) aff. crosbeyi Amellago I Shells
(Orthodactylites) cf. semiannulatum Amellago I Shells
(Orthodactylites) andaluciensis Agounou I Shells
sp. Ait El Abbas I Shells
?Dicoelites[33] sp. A Iwansitn III Rostra Dicoelitidae
Discohelix[31] subaequalis Rich region; Outarbat III-IV Shells Discohelicidae
pulchrior Rich region IV Shells
sp. Rich region IV Shells
Docidoceras[1] wysogorskii Rich III Shells Otoitidae
Eleganticeras[34] exaratum Amellago I Shells Dactylioceratidae
sp. Amellago I Shells
Emileia[35] (Emileia) polymera Boukendil, Assameur IV Shells Otoitidae
Eucyclus[31] ornatus Rich region IV Shells Eucyclidae
sp. Aberdouz III Shells
Eucycloidea[31] (Pseudalaria?) subangulata Rich region IV Shells Eucyclidae
cf. etheridgii Rich region; Outarbat III-IV Shells
Eudmetoceras[1] flexuosum Rich II–III Shells Hammatoceratidae
masticonense Rich III Shells
eudmetum Rich III Shells
sp. Rich II–III Shells
Euaptetoceras[1] amaltheiforme Rich III Shells Hammatoceratidae
amplectens Rich III Shells
sp. Rich II–III Shells
Fontannesia[1] aurita Rich II Shells Sonniniidae
montillanensis Rich II Shells
sp. Rich II–III Shells
Furloceras[30] cf.chelussii Ait El Abbas I Shells Phymatoceratidae
Geczyceras[30] clausum Ait El Abbas I Shells Hammatoceratidae
speciosum Azag I Shells
Grammatodon[1] sp. Rich II–III Shells Parallelodontidae
Hammatoceras[34] H. aff. insigne n’Zala (Jbel Aouja) I Shells Hammatoceratidae
Harpoceras[30][34] falciferum Amellago I Shells Hildoceratidae
pseudoserpentinum Amellago I Shells
serpentinum Amellago I Shells
subplanatum Amellago I Shells
cf.mediterraneum Azag I Shells
Hildaites[30][34] forte Amellago I Shells Hildoceratidae
levisoni Amellago I Shells
cf. serpentiniformis Amellago I Shells
cf. subserpentinus Amellago; Agounou I Shells
striatus Amellago I Shells
wrighti Amellago I Shells
(Martanites) cf. prorsiradiatus Agounou I Shells
murleyi Agounou I Shells
aff. murleyi Ait El Abbes I Shells
cf. striatus Agounou I Shells
undicosta Agounou I Shells
Hildoceras[21][30][34] bifrons Amellago; Azag; Talghemt Ridge I Shells Hildoceratidae
lusitanicum Amellago; Talghemt Ridge I Shells
semipolitum Amellago; Talghemt Ridge I Shells
sublevisoni Amellago; Azag; Talghemt Ridge I Shells
bonarellii Talghemt Ridge I–II Shells
cf.tethysi Azag I Shells
appenninicus Agounou; Azag I Shells
snoussi Ait El Abbes I Shells
cf.apertum Azag I Shells
sp. Amellago; Azag I Shells
Hyperlioceras[1] walkeri Rich III Shells Graphoceratidae
incisum Rich III Shells
inclusum Rich III Shells
mundum Rich III Shells
sp. Rich III Shells
Leioceras[32] cf. comptum n’Zala (Jbel Aouja) II Shells Graphoceratidae
costusum n’Zala (Jbel Aouja) II Shells
Lithophaga[17] sp. Boukendil area IV Shells, borings Mytilidae
Ludwigia[32] aff. haugi n’Zala (Jbel Aouja) II Shells Graphoceratidae
Lytoceras[30][34][36] siemensi Amellago; n’Zala I Shells Lytoceratidae
cf.siemensi Agounou; Ait El Abbes I-II Shells
subfimbriatum n’Zala II Shells
francisci Ait El Abbes I-II Shells
sp. Amellago; Agounou; Ait El Abbes; n’Zala I–II Shells
Maconiceras[34] soloniacense Amellago I Shells Hammatoceratidae
Malladaites[1] pertinax Rich III Shells Hammatoceratidae
Merlaites[30] alticarinatus Ait El Abbes; Azag I Shells Hildoceratidae
gradatus Ait El Abbes I Shells
Neotelodactylites[30] eucosmus Ait El Abbes I Shells Dactylioceratidae
Nerinea[17] sp. Assameur n’Ait Fergane IV Shells Nerineidae
Nicaniella[1] sp. Rich, Amellago II–III Shells Astartidae
Nodicoeloceras[30] cf. merlai Ait El Abbes I Shells Dactylioceratidae
Osperleioceras[30] bicarinatum Ait El Abbes; Azag I Shells Hildoceratidae
beauliziense Azag I Shells
cf. authelini Azag I Shells
cf. reynesi Azag I Shells
O. (Pseudolioceras) sp. Ait El Abbes I Shells
sp. Azag I Shells
Palaeonucula[1] hammeri Rich, Amellago II–III Shells Nuculidae
Palaeoneilo[1] sp. Rich III Shells Malletiidae
Peronoceras[30] cf. subarmatum Ait El Abbes I Shells Dactylioceratidae
Phylloceras[30][34] doderleinianum Ait El Abbes II Shells Phylloceratidae
sp. Amellago; n’Zala I–II Shells
Phymatoceras[30] sp. Azag I Shells Phymatoceratidae
Pholadomya[27] (Bucardiomya) bucardium Jebel Assameur; Boukendil; Rich-Gourrama area IV Shells Pholadomyidae
Planammatoceras[32] sp. n’Zala (Jbel Aouja) II Shells Hammatoceratidae
Platystrophites[30] cf. latus Agounou I Shells Dactylioceratidae
Pleurotomaria[31] armata Rich region IV Shells Pleurotomariidae
Pleydellia[32] aalensis n’Zala (Jbel Aouja) II Shells Graphoceratidae
Polyplectus[34] pluricostatum Amellago I Shells Hammatoceratidae
sp. Amellago I Shells
Porpoceras[30][34] gr. vortex-verticosum Amellago; Talghemt Ridge I Shells Dactylioceratidae
cf. vortex Ait El Abbes I Shells
cf. vorticellum Ait El Abbes I Shells
cf. verticosum Ait El Abbes I Shells
sp. Ait El Abbes; Agounou I Shells
Praestrigites[1] sp. Rich III Shells Strigoceratidae
Praeoppelia[1] sp. Rich III Shells Oppeliidae
Protetragonites[36] sp. n’Zala II Shells Lytoceratidae
Protoecotraustes[1] sp. Rich III Shells Haploceratidae
Pseudocrassiceras[30] aff.bayani Ait El Abbes; Azag I Shells Hildoceratidae
bayani Ait El Abbes I Shells
frantzi Ait El Abbes I Shells
praegruneri Ait El Abbes I Shells
cf. frantzi Azag I Shells
sp. Azag I Shells
Pseudogrammoceras[30] praegruneri Azag I Shells Hildoceratidae
subregale Ait El Abbes; Azag I Shells
aff. fallaciosum Azag I Shells
Rastellum[17] sp. Assameur n’Ait Fergane, Boukendil IV Shells Ostreidae
Rhynchocerithium[31] sp. Rich region IV Shells Procerithiidae
Riselloidea[31] sp. Outarbat III-IV Shells Eucyclidae
Sadkia[31] richensis Rich region IV Shells Eucyclidae
Saxicava[17] sp. Boukendil area IV Shells, borings Hiatellidae
Sonninia[1] dominans Rich III Shells Sonniniidae
modestum Rich III Shells
nannomorpha Rich III Shells
Tmetoceras[32] T. scissum n’Zala (Jbel Aouja) I–II Shells Hammatoceratidae
Toxamblyites[35] densicostatus Boukendil, Assameur IV Shells Haploceratidae
Trigonia[1] sp. Rich III Shells Trigoniidae
Trilobiticeras[1] platygaster Rich III Shells Otoitidae
trilobitoides Rich III Shells
sp. Rich III Shells
Tylostoma[1] globosum Rich III Shells Tylostomatidae
Vacekia[32] sp. n’Zala (Jbel Aouja) I–II Shells Graphoceratidae
Zugodactylites[30] sp. Ait El Abbes I Shells Dactylioceratidae
Zurcheria[1] sp. Rich III Shells Hammatoceratidae

Arthropods

Taxa Species Locality Member Material Notes Images
Bairdia[21] sp. A Talghemt Ridge I–II Valves Bairdiidae
aff. hilda Talghemt Ridge IV Valves Bairdiidae
Bairdiacypris[21][22] sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge I–II Valves Bairdiacyprididae
Cytherella[21][22] aff. concentrica Talghemt Ridge IV Valves Cytherellidae
aff. fullonica Talghemt Ridge IV Valves Cytherellidae
aff. toarcensis Talghemt Ridge I–II Valves Cytherellidae
spp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge II Valves Cytherellidae
Cytherelloidea[21] gr. catenulata Talghemt Ridge IV Valves Cytherelloididae
Favreina[23] isp. Assameur, Jebel Taferdout III-IV Coprolites Decapod Coprolites (Axiidea?)
Kinkelinella[21] K. sermoisensis Talghemt Ridge I–II Valves Kinkelinellidae
gr. sermoisensis Talghemt Ridge I–II Valves Kinkelinellidae
Micropneumatocythere[22] sp. Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge II Valves Podocopida
Monoceratina[21] aff. ventrale Talghemt Ridge IV Valves Monoceratidae
Paracypris[21] sp. Talghemt Ridge IV Valves Cytheridae
Polycope[21] sp. Talghemt Ridge IV Valves Polycopidae
Pontocyprella[21] sp. Talghemt Ridge I–II Valves Cyprididae
Procytheridea[21][22] sp. B Assameur n’Aït Fergane; Bou Kendill; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi; Talghemt Ridge II-IV Valves Cytherideidae
Striatojonesia[21] S. striata Talghemt Ridge IV Valves Bythocytheridae

Echinoderms

Genus Species Locality Member Material Notes Images
Cidaris[27] sp. Jebel Assameur; Boukendil; Rich-Gourrama area IV Spines & Specimens Cidaridae
Diplopodia[22][27] cf. morierii Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi IV Spines & Specimens Diplopodiidae
Gymnocidaris[27] pustulosa Jebel Assameur; Boukendil; Rich-Gourrama area IV Spines & Specimens Hemicidaridae
Hemicidaris[22][27] koechlini Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi IV Spines & Specimens Hemicidaridae
Psephechinus[22][27] globosus Boukendil; Rich-Gourrama area; Assameur n’Aït Fergane; Jbel Tazigzaout; Ifrda n’Tizguienne; Assameur n’Baknou-Aferdoun’Achi IV Spines & Specimens Stomechinidae

Annelida

Genus Species Locality Member Material Notes Images
Nogrobs[35] moroccensis Boukendil, Assameur IV Quadrangular tubes Serpulidae
Serpula[9][13][17] socialis Assameur n’Ait Fergane, Boukendil; Imilchil; Jebel Bou Kendill (cap bed) I-III–IV Tubes / colonies Serpulidae
sp. Assameur n’Ait Fergane, Boukendil; Jebel Bou Kendill (cap bed) III–IV Tubes / colonies Serpulidae
Terebella[17] lapilloides Assameur n’Ait Fergane, Boukendil; Jebel Bou Kendill (cap bed) III–IV Tubes / colonies Terebellidae

Chondrichthyes

Taxa Species Locality Member Material Notes Images
Strophodus[37] atlasensis Talsint (eastern High Atlas) III–IV Articulated dentition (holotype PIMUZ A/I 5181) Acrodontidae

Dinoflajellates

Taxa Species Locality Member Material Notes Images
Kallosphaeridium[38] sp. Amellago I-II Cysts Incertade Sedis
Luehndea[38] cirilliae Amellago I Cysts Luehndeoideae
spinosa Amellago I Cysts Luehndeoideae
Mancodinium[38] semitabulatum Amellago I-III Cysts Mancodinioideae
Mendicodinium[38] microscabratum Amellago I-II Cysts Gonyaulacales
sp. 1 Amellago I-II Cysts Gonyaulacales
Moesiodinium[38] sp. Amellago I-II Cysts Heterocapsaceae
Phallocysta[38] sp. Amellago I-II Cysts Peridiniphycideae
Wallodinium[38] cf. cylindricum Amellago I-II Cysts Incertade Sedis

Algae

Taxa Species Locality Member Material Notes Images
Biscutum[10][39][40] depravatum Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
grande Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
Botryococcus[38] sp. Amellago I-III Miospore Botryococcaceae
Cayeuxia[17][18] sp. Assameur n’Ait Fergane; Assoul ramp IV Filamentous thalli Cyanobacteria
Carinolithus[10][39][40] superbus Amellago; Talghemt Ridge II–III Coccoliths Carinolithaceae
Cylindroporella[17] arabica Assameur n’Ait Fergane IV Thalli Dasycladaceae
Discorhabdus[10][39][40] ignotus Amellago; Talghemt Ridge II–III Coccoliths Discorhabdaceae
striatus Amellago; Talghemt Ridge II–III Coccoliths Discorhabdaceae
Garwoodia[20] sp. Amellago canyon IV Filamentous thalli Cyanobacteria
Girvanella[17] sp. Assameur n’Ait Fergane IV Filamentous thalli Cyanobacteria
Halimeda[17] sp. (cf.) Assameur n’Ait Fergane IV Thallus fragments Halimedaceae
Lotharingius[10][39][40] hauffii Amellago; Talghemt Ridge II–III Coccoliths Lotharingiaceae
Mitrolithus[10][39][40] jansae Amellago; Talghemt Ridge II–III Coccoliths Mitrolithaceae
Permocalculus[17] sp. Assameur n’Ait Fergane, Boukendil IV Encrusting thalli Gymnocodiaceae
Rivularia[20] sp. Amellago canyon IV Filamentous thalli Rivulariaceae
Sarfatiella[17] dubari Assameur n’Ait Fergane IV Thalli Dasycladaceae
Schizosphaerella[10][39][40] sp. Amellago; Talghemt Ridge II–III Coccoliths Schizosphaerellaceae
Sellinoporella[17] sp. Assameur n’Ait Fergane IV Thalli Dasycladaceae
Similiscutum[10][39][40] finchii Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
novum Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
Solenopora[17] sp. (cf.) Assameur n’Ait Fergane IV Encrusting thalli Solenoporaceae
Tasmanites[38] sp. Amellago I-III Miospore Tasmanaceae
Triscutum[10][39][40] sullivanii Amellago; Talghemt Ridge II–III Coccoliths Biscutaceae
Watznaueria[10][39][40] colacicchii Amellago; Talghemt Ridge III Coccoliths Watznaueriaceae
fossacincta Amellago; Talghemt Ridge III Coccoliths Watznaueriaceae

Plants

Taxa Species Locality Member Material Notes Images
Alisporites[38] spp. Amellago I Pollen Corystospermaceae
Callialasporites[38] spp. Amellago I-II Pollen Araucariaceae
Classopollis[38] sp. Amellago I Pollen Hirmeriellaceae
Cyathidites[38] spp. Amellago I Spores Cyatheaceae
Deltoidospora[38] spp. Amellago I Spores Pteridophyte
Kraeuselisporites[38] reissingeri Amellago I Spores Lycopodiaceae
Ischyosporites[38] variegatus Amellago I Spores Lycopodiaceae
Neoraistrickia[38] spp. Amellago I-II Spores Selaginellaceae
Quadraeculina[38] spp. Amellago I Pollen Podocarpaceae
Zonalapollenites[38] spp. Amellago I-II Pollen Araucariaceae

See also

References

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